Department of Earth and Environmental Sciences
Newton Horace Winchell School of Earth and Environmental Sciences

Robert A. Martin


Contact Information

Home: 803 Marshall Dr., Erie PA 16505

Phone: 270/978-0049



Professional preparation

University of Florida      Gainesville, FL             Zoology                                      PhD 1969

Tulane University          New Orleans, LA          Biology                                       MS   1967

Hofstra University         Hempstead, NY            Biology                                       BA   1965



2016-                         Adjunct Professor, Dept. of Earth Sciences, Univ. of Minnesota

2013-                         Professor Emeritus, Murray State University

1993-2013                 Professor, Dept. of Biology, Murray State University

1986-1993                 Professor, Dept. of Biology, Berry College

1972-1986                 Asst. Prof. – Prof., Fairleigh Dickinson University

1969-1972                 Asst. Professor, Dept. of Biology, South Dakota School Mines & Technology


 Select Publications 


     _________,  P. Peláez-Campomanes, L. Viriot. 2017. First report of rodents from the late Hemphillian (late Miocene) Zwiebel Channel    and a revised late Neogene biostratigraphy/biochronology of the Sand Draw area of Nebraska. Historical Biology pdf


  • __________. 2016. Geomys tyrioni, a new species of early Pleistocene dwarf pocket gopher from the Meade Basin of southwestern Kansas. Journal of Mammalogy DOI: 10.1093/mammal/gyw024. pdf
  • __________. 2016. Body size in (mostly) mammals: mass, speciation rates, and the translation of gamma to alpha diversity on evolutionary timescales. Historical Biology.
  • __________ and P. Peláez-Campomanes. 2016. Extinction Rates of the Meade Basin Rodents: Application to Current Biodiversity Losses. Lethaia (in press).


  • __________. 2015. A brief review of the Spanish archaic Pleistocene arhizodont voles; Pp. 207-226, in L. Gibert, C. Ferrández-Cañadell (eds.). Geology and Paleontology of Cueva Victoria. Mastia, Nos. 11-13: Archeological Museum of Cartagena, Spain. pdf

  • L. Gibert, G. R. Scott, D. Scholz, A. Budsky, C. Ferrandez, _____________, F. Ribot, M. Lería. 2015. Chronology for the Cueva Victoria fossil site (SE Spain): evidence for Early Pleistocene Afro-Iberian dispersal.  Journal of Human Evolution pdf
  • Mutttoni, G.,  D. V. Kent, G. Scardia and __________. 2015. Bottleneck at Jaramillo for human migration to Iberia and rest of Europe? Journal of Human Evolution 80:187-190. pdf

  • ___________. 2015. A preliminary diversity-divergence model for arvicolid rodents. Paleobiodiversity and Palaeoenvironments, 95: 253-256. pdf

  • Fox, D. L., ______________, and others. 2015. Biotic and abiotic forcing during the transition to modern grassland ecosystems: evolutionary and ecological responses of small mammal communities over the last 5 million years. Pp. 197-216 In Polly, P. D., J. J. Head and D. L. Fox (eds.). Earth-Life Transitions: Paleobiology in the Context of Earth System Evolution. Paleontological Society Papers 21.


  • __________ and P. Peláez-Campomanes. 2014. Diversity dynamics of the late Cenozoic rodent community from southwestern Kansas: the influence of historical processes on community structure. Journal of Quaternary Science 29:221-23. pdf
  • ________. 2014. A critique of vole clocks. Journal of Quaternary Science Reviews. 94:1-6. pdf

  • __________ and K. O’Bryan. 2014. Size and shape variation in late Pleistocene pine vole (Mammalia: Arvicolidae: Pitymys pinetorum) first lower molars from three caves in Kentucky and Georgia. Paludicola 9:159-166. pdf

  • ________. 2014. Late Neogene beaver Dipoides from the Pipe Creek Sinkhole of Grant County, Indiana. Paludicola 9:167-175. pdf

  • ________. 2014. Missing Links: Evolutionary Concepts & Transitions Through Time. McDonald & Woodward, Granville, OH. 434 p.


  • Fox, D. L., J. G. Honey, _________, and P. Peláez-Campomanes. 2012a. Pedogenic carbonate stable isotope record of environmental change during the Neogene in the southern Great Plains, southwest Kansas, USA: Oxygen isotopes and paleoclimate during the evolution of C4-dominated grasslands. GSA Bulletin 124:432-443. pdf
  • Fox, D. L., J. G. Honey, _________, and P. Peláez-Campomanes. 2012b. Pedogenic carbonate stable isotope record of environmental change during the Neogene in the southern Great Plains, southwest Kansas, USA: Carbon isotopes and the evolution of C4-dominated grasslands. GSA Bulletin 124:444-462. pdf
  • __________,  P. Peláez-Campomanes and C. Mecklin. 2012. Patterns of size change in late Neogene pocket gophers from the Meade Basin of Kansas and Oklahoma. Historical Biology 24:537-545. pdf
  • ________. 2012.Victoriamys, a new generic name for Chaline’s vole from the Pleistocene of western Europe. Geobios 45:445-450. pdf


  • ___________, P. Peláez-Campomanes, and J. G. Honey. 2011. Preliminary study of rodents from the late Pleistocene Golliher B assemblage of Meade County, Kansas indicates an intense cold period near the end of the Pleistocene.  Palaeontologica Electronica 14.3.35A: 1-13. pdf
  • ___________, P. Peláez-Campomanes, J. G. Honey, F. Marcolini and W. A. Akersten. 2011. Five million years of pocket gopher history in the Meade Basin of southwestern Kansas and northwestern Oklahoma. Journal of Vertebrate Paleontology 31: 866-884. pdf


  • __________. 2010. The North American Promimomys immigration event. Paludicola 8:14-21. pdf


  • Marcolini, F., P. Piras and _______________. 2009. Testing evolutionary dynamics on first lower molars of Pliocene Ogmodontomys (Arvicolidae, Rodentia) from the Meade Basin of southwestern Kansas (USA): a landmark-based approach. Palaios 24:535-543. pdf
  • ___________, F. Marcolini and F. Grady. 2009. The early Pleistocene Hamilton Cave muskrats and a review of muskrat size change through the late Neogene. Paludicola 7:61-66. pdf


  • __________,  P. Peláez-Campomanes, J. G. Honey, D. L. Fox, R. J. Zakrzewski, L. B. Albright, E. H. Lindsay, N. D. Opdyke, and H. T. Goodwin. 2008. Rodent community change at the Pliocene-Pleistocene transition in southwestern Kansas and identification of the Microtus immigration event on the Central Great Plains. Palaeogeography, Palaeoclimatology, Palaeoecology 267:196-207. pdf
  • Marcolini. F. and ___________. 2008. Mosaic evolution in first lower molars of Pliocene Ogmodontomys (Rodentia: Arvicolidae) from the Meade Basin of southwestern Kansas (USA). N. Jarbuch f. Geol. und Palaontol. 249:313-342. pdf
  • ___________, A. Siefker and F. Marcolini. 2008. Modeling the morphology and evolution of the linea sinuosa (crown-root junction) in arvicolid rodents; a test with Pliocene Ogmodontomys from Kansas, USA. Lethaia 42:155-166. pdf


  • L. Gibert, G. Scott, ____________, and J. Gibert. 2007. The early to middle Pleistocene boundary in the Baza Basin (Spain). Quaternary Science Reviews  26: 2067-2089. pdf

  • ____________. 2007. Arvicolidae; pp. 480-497 in Janis, C. M., G. G. Gunnell, and L. Jacobs eds.), Evolution of Tertiary Mammals of North America. Volume II.Cambridge, New York. pdf


  • ____________, C. P. Crockett and F. Marcolini. 2006. Variation of the schmelzmuster and other enamel characters in the primitive Pliocene vole Ogmodontomys from Kansas. Journal of Mammalian Evolution 13: 223-241. pdf


  • Honey, J. G., Peláez-Campomanes and ______________. 2005. Stratigraphic framework of early Pliocene fossil localities along the north bank of the Cimarron River, Meade County, Kansas. Ameghiniana 42: 461-472. pdf
  • Peláez-Campomanes and ________________. 2005. The Pliocene and Pleistocene history of cotton rats in the Meade Basin of southwestern Kansas. Journal of Mammalogy 86: 469-484. pdf
  • ___________. 2005. Preliminary correlation of Florida and Central Great Plains Pliocene and Pleistocene mammalian local faunas based on rodent biostratigraphy; pp. 363-367 in G. Morgan and R. Hulbert (eds.), Bulletin Florida State Museum 45.


  • ____________, R. T. Hurt, J. G. Honey, and P. Peláez-Campomanes. 2003. Late Pliocene and early Pleistocene rodents from the northern Borchers Badlands (Meade County, Kansas), with comments on the Blancan-Irvingtonian boundary in the Meade Basin. Journal of Paleontology 77:985-1001. pdf
  • ____________. 2003. Biochronology of Pliocene and Pleistocene arvicolid rodents from the central Great Plains of North America. Colloquios de Paleontologia, University Complutense, Madrid  Vol. Ext. I:373-383. pdf
  • __________, L. Duobinis-Gray, and C. P. Crockett. 2003. A new species of early Pleistocene Synaptomys and its relevance to southern bog lemming origins. Journal of Vertebrate Paleontology 23: 917-936. pdf  

  • __________. 2003. The status of Mimomys in North America revisited. Deinsea 10: 399-406.


  • ____________, J. G. Honey, P. Peláez-Campomanes, H. T. Goodwin, J. A. Baskin, and R. J. Zakrzewski. 2002. Blancan lagomorphs and rodents of the Deer Park assemblages, Meade County, Kansas. Journal of Paleontology 76:1072-1090. pdf

  • ___________, H. T. Goodwin, and J. O. Farlow. 2002. Late Neogene rodents from the Pipecreek Sinkhole, Grant County, Indiana. Journal of Vertebrate Paleontology 22:137-151. pdf

  • ___________. 2002. A possible role for reproductive behavior in the origin of tetrapod feet. Paludicola 4:6-9.


  • ___________. 2001. A preliminary list of late Pleistocene mammals from the Peccary Room, Yarbrough Cave, Bartow County, Georgia. Paludicola 3:33-39.


  • ____________. 1996. Tracking mammal body size distributions in the fossil record: a preliminary test of the 'rule of limiting similarity.' Acta Zool.Cracov. 39: 321-328. pdf

  • ____________. 1996. Dental evolution and size change in the North American muskrat: classification and tempo of a presumed phyletic sequence; pp 431- 457 in  Palaeoecology and Palaeonvironments of Late Cenozoic Mammals. K. Seymour and K. Stewart (eds.)., Univ. Toronto Press, Canada.

  • ____________, and A. D. Barnosky (eds.). 1993. Morphological Change in Quaternary Mammals of North America. Cambridge U. Press, New York. 415 p. 

  • ____________. 1993. Patterns of variation and speciation in Quaternary rodents; pp.226-280. In  Morphological Change in Quaternary Mammals of North America. R. A. Martin and A.D. Barnosky (eds.). Cambridge U. Press, New York.

  • ____________. 1992. Generic species richness and body mass in North American mammals: support for the inverse relationship of body size and speciation rate. Histor. Biology 6:73-90. pdf

  • ____________. 1991. Evolutionary relationships and biogeography of late Pleistocene prairie voles from the eastern United States. Scientific Papers, Illinois State Museum 23: 251-260.

  • ____________,   and R.H. Prince. 1990. Variation and evolutionary trends in the dentition of late Pleistocene Microtus pennsylvanicus  from three levels in Bell Cave, Alabama. Histor. Biology 4: 117-129. 

  • ___________. 1990. Estimating body mass and correlated variables in extinct mammals:  travels in the fourth dimension. Pp. 59-68 in  Body Size in Mammalian Paleobiology, J. Damuth and B. MacFadden (eds), Cambridge Univ. Press, New York. 

  • ____________. 1989. Early Pleistocene zapodid rodents from the Java local fauna of north-central South Dakota.  J. Vert. Paleontol. 9: 101-109 pdf

  • ____________. 1986. Energy, ecology and cotton rat evolution.  Paleobiology, 12: 370-382. pdf

  •    ____________. 1980. Body mass and basal metabolism of extinct mammals.  Comp Biochem. Physiol., 66A: 307-314. pdf